Asian forest tortoise

Asian forest tortoise
Juvenile mountain tortoise, Manouria emys
Conservation status
Endangered (IUCN 3.1)
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Family:	Testudinidae
Genus:	Manouria
Species:	M. emys
Binomial name
Manouria emys (Schlegel & Müller, 1844)
Synonyms
Manouria emys emys Testudo emys Schlegel & Müller, 1840 Testudo emydoides Duméril & Bibron, 1851 Manouria fusca Gray, 1854 Teleopus luxatus LeConte, 1854 Manouria emydoides Strauch, 1862 Manouria emys Strauch, 1862 Manouria luxata Strauch, 1862 Testudo (Scapia) falconeri Gray, 1869 Scapia falconeri Gray, 1870 Manuria emys Lydekker, 1889 Geochelone emys Loveridge & Williams, 1957 Manouria emys emys Obst, 1983 Geochelone emys emys Gosławski & Hryniewicz, 1993 Testudo emys emys Paull, 1999 Manouria emys phayrei Testudo phayrei Blyth, 1853 Testudo (Scapia) falconeri Gray, 1869 Scapia falconeri Gray, 1870 Scapia phayrei Gray, 1871 Testudo nutapundi Reimann, 1979 Geochelone nutapundi Groombridge, 1982 Manouria emys nutapundi Obst, 1983 Manouria emys phayrei Bour, 1984 Geochelone emys nutapundi Gosławski & Hryniewicz, 1993 Manouria nutapundi Obst, 1996 Manouria emys phayeri Paull, 1997 Manouria emys phayre Das, 2001 (ex errore) Manouria emys phareyi Ferri, 2002 (ex errore)

Asian forest tortoise (Manouria emys), also known as Asian brown tortoise, is a species of tortoise found in India (Assam), Bangladesh, Burma (or Myanmar), Thailand, Malaysia and Indonesia (Sumatra, Borneo).

Description

The largest tortoise in mainland Asia; the large adult of the northern subspecies, Manouria emys phayrei, can reach 25 kg in the wild and much more than that in captivity. Shell considerably depressed, its depth not half its length; anterior and posterior margins reverted, more or less strongly serrated; nuchal present; supracaudal shields two; dorsal shields concentrically striated, often concave; vertebrals much broader than long and at least as broad as costals. Plastron large, gular region somewhat produced and usually notched, hind lobe deeply notched; the pectoral shields may be widely separated from each other, or from a short median suture; axillary shield very small, inguinal large. Head moderate; two large prefrontal shields and a large frontal; beak not hooked; jaws feebly denticulated, the alveolar surface of the upper jaw with a strong median ridge. Forelimb anteriorly with very large, bony, pointed, imbricate tubercles, forming four or five longitudinal series; hind limb with very large bony tubercles on the plantar surface, with others larger, conical, and spur-like on the heel, and a group of still larger conical tubercles on each side on the back of the thighs. Adult dark brown or blackish; carapace of young yellowish brown, with dark-brown markings.

Believed to be among the most primitive of living tortoises, based on molecular and morphological studies. This is the only tortoise which lays its eggs above ground in a nest, which the female constructs of leaf litter. The female uses both front and rear legs to gather material for the nest and lays up to 50 eggs deep inside it. She then sits on and near the nest to protect it, and will 'chase' predators and intruders away.

Subspecies

• Manouria emys emys: S Thailand, Malaysia, Sumatra, Borneo; Type locality: Sumatra. M. e. emys has separated pectoral scutes while M. e. phayrei has joined pectoral scutes.
• Manouria emys phayrei: N/W Thailand to NE India; Type locality: Arakan; Tenasserim Provinces'. M. e. phayrei has been named after Sir Arthur Purves Phayre (1812–1885), British Army officer in India who became Commissioner of British Burma.

Other references

• Asian Turtle Trade Working Group (2000). Manouria emys. 2006. IUCN Red List of Threatened Species. IUCN 2006. www.iucnredlist.org. Retrieved on 11 May 2006. Listed as Endangered (EN A1cd+2 cd v2.3)
• Anderson, J. 1871 On Testudo Phayrei, Theob. & Dr. Gray. Ann. Mag. Nat. Hist. (4) 8: 324-330
• Blyth, E. 1854 Notices and descriptions of various reptiles, new or little-known. Part I. J. Asiat. Soc. Bengal 22 [1853]: 639-655
• Schlegel, H. & Müller, S. 1844 Over de Schildpadden van den Indischen Archipel. In: Temminck,V. (1839–1847) Verhandelingen over de natuurlijke geschiedenis der Nederlandsche overzeesche bezittingen, door de leden der Natuurkundige Commisie in Oost-Indie en andere schrijvers. Leijden folio. Afd. 1 Zoologie in 12 afleveringen, met 45 gekl. pl. - Vitg. door C.J. Temminck, Leiden, Leuchtmans u. Hoeck in comm.: 29-36
• Stoliczka, F. 1871 Note on Testudo Phayrei. Ann. Mag. Nat. Hist. (4) 8: 212

External links

• Manouria emys at the Reptarium.cz Reptile Database
• http://www.tortoise.org/archives/manouria.html