Goniopholididae

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Goniopholidids
Temporal range: 196–65 Ma
Sinemurian - Maastrichtian
250px
Dollo’s goniopholidid (Unnamed specimen IRSNB R47), Musee d'Histoire Naturelle, Brussels
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Superorder: Crocodylomorpha
Clade: Neosuchia
Family: Goniopholididae
Cope, 1875
Subgroups

See text.

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Goniopholididae is an extinct family of moderate-sized semi-aquatic crocodyliforms superficially similar to living crocodiles. They lived between the Early Jurassic and the Late Cretaceous.[1][2]

Phylogeny

The following cladogram simplified after an analysis presented by Marco Brandalise de Andrade and colleagues in 2011.[3]

Neosuchia
Atoposauridae

Theriosuchus pusillus



Theriosuchus guimarotae






Rugosuchus




Bernissartia



Eusuchia






Stolokrosuchus





Tethysuchia



Thalattosuchia



Goniopholididae

Calsoyasuchus valliceps





"Goniopholis" phuwiangensis




Eutretauranosuchus delfi



"Sunosuchus" junggarensis





Sunosuchus miaoi



Chalawan thailandicus






Siamosuchus phuphokensis




Amphicotylus lucasii




Denazinosuchus kirtlandicus



Nannosuchus gracilidens





Hulkepholis willetti




Anteophthalmosuchus



Dollo’s goniopholidid




Goniopholis

Goniopholis baryglyphaeus




Goniopholis kiplingi



Goniopholis simus














Geographical distribution

Goniopholidids are known across Laurasia, ranging from North America, Europe and China from the Middle Jurassic,[1][4] and reaching Thailand by the Early Cretaceous.[5]

Biology

Compared to modern crocodilians, goniopholids are very unusual in several respects. They possessed two rows of rectangular, interlocking osteoderms like those of terrestrial crocodilymorphs like atoposaurids, that are relatively simple, do not extend far in their necks, as opposed to the ornate armours of modern crocodilians; likewise, unlike modern crocodilians but like many extinct forms like phytosaurs, they have ventral osteoderms as well. Their forelimbs are also proportionally very long, particularly in the humeri and wrist bones, being as long or longer than the hindlimbs, the opposite of the condition seen in modern crocodilians. Some like Anteophthalmosuchus also have forwardly oriented eyes, as opposed to the dorsally oriented eyes seen in modern forms. These suggest multiple biomechanical differences from modern species.[6][7]

References

  1. 1.0 1.1 Steel R. 1973. Crocodylia. Handbuch der Paläoherpetologie, Teil 16. Stuttgart: Gustav Fischer Verlag, 116 pp.
  2. Tykoski RS, Rowe TB, Ketcham RA, Colbert MW. 2002. Calsoyasuchus valliceps, a new crocodyliform from the Early Jurassic Kayenta Formation of Arizona. Journal of Vertebrate Paleontology 22 (3): 593-611.
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  4. Maisch MW, Matzke AT, Stohr H. 2003. Sunosuchus (Archosauria, Crocodyliformes) from the Toutunhe Formation (Middle Jurassic) of the Southern Junggar Basin (Xinjiang, NW-China). Geobios 36 (4): 391-400.
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  6. Salisbury, S. W. & Frey, E. 2000. A biomechanical transformation model for the evolution of semi-spheroidal articulations between adjoining vertebral bodies in crocodilians. In Grigg, G. C., Seebacher, F. & Franklin, C. E. (eds) Crocodilian Biology and Evolution. Surry Beatty & Sons (Chipping Norton, Aus.), pp. 85-134.
  7. Salisbury, S. W. & Naish, D. (2011). Crocodilians. In Batten, D. J. (ed.) English Wealden Fossils. The Palaeontological Association (London), pp. 305-369.

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