2013 in paleontology

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Paleontology, palaeontology or palæontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because mankind has encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2013.

Plants

Angiosperms

Name Novelty Status Authors Age Unit Location Notes Images

Atli[2]

Gen. et sp. nov.

Valid

Smith et al.

Late Cretaceous (Campanian)

 Canada

A ranunculalean known from a stem with lianoid anatomy. The type species is Atli morinii.

Bambusiculmus[3]

Gen. et 2 sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

 China

A bamboo. The type species is Bambusiculmus angustus; genus also includes Bambusiculmus latus.

Bambusium angustifolia[3]

Sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

 China

A bamboo, a species of Bambusium.

Bambusium latifolia[3]

Sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

 China

A bamboo, a species of Bambusium.

Bombacacioxylon[4]

Gen. et sp. nov.

Valid

Mehrotra et al.

Late Miocene to early Pliocene

Tipam Group

 India

A possible relative of Bombax. The type species is Bombacacioxylon tertiarum.

Firmiana oligocenica[5]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Firmiana.

Guadua morronei[6]

Sp. nov.

Valid

Brea, Zucol & Franco

Pliocene–Pleistocene

Ituzaingó Formation

 Argentina

A bamboo, a species of Guadua.

Iterophyllum[7]

Gen. et sp. nov.

Valid

Barral et al.

Early Cretaceous (late Barremian)

 Spain

A basal eudicot. The type species is Iterophyllum lobatum.

Jaguariba wiersemana[8]

Gen. nov. et sp. nov.

Valid

Coiffard, Mohr & Bernardes-de-Oliveira

Early Cretaceous

Crato Formation

 Brazil

Fossil Nymphaeales.

Palmoxylon vastanensis[9]

Sp. nov.

Valid

Prasad et al.

Eocene

Cambay Shale Formation

 India

A palm tree, a species of Palmoxylon.

Paranephelium makumensis[5]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Paranephelium.

Poeciloneuron preindicum[10]

Sp. nov.

Valid

Srivastava & Mehrotra

Late Oligocene

 India

A species of Poeciloneuron.

Potomacapnos[11]

Gen. et sp. nov.

Valid

Jud & Hickey

Early Cretaceous (Aptian)

Potomac Group

 United States

A close relative of ranunculalean eudicots. The type species is Potomacapnos apeleutheron.

Pterygota palaeoalata[5]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Pterygota.

Sapindus palaeoemarginatus[5]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Sapindus.

Spixiarum kipea[12]

Gen. nov. et sp. nov.

Valid

Coiffard, Mohr & Bernardes-de-Oliveira

Early Cretaceous

Crato Formation

 Brazil

Fossil Aroid.

Virola dominicana[13]

Sp. nov.

Valid

Poinar & Steeves

Cenozoic (Eocene or Miocene)

 Dominican Republic

A member of Myristicaceae found in Dominican amber, a species of Virola.

Other seed plants

Name Novelty Status Authors Age Unit Location Notes Images

Bancroftiastrobus[14]

Gen. et sp. nov.

Valid

Rothwell et al.

Late Jurassic (early Kimmeridgian)

Kimmeridge Clay Formation

 United Kingdom

A member of Coniferales of uncertain phylogenetic placement. The type species is Bancroftiastrobus digitata.

Chengia[15]

Gen. et sp. nov.

Valid

Yang, Lin & Wang

Early Cretaceous (Aptian)

Yixian Formation

 China

A member of Ephedraceae. The type species is Chengia laxispicata.

Cunninghamia taylorii[16]

Sp. nov.

Valid

Serbet, Bomfleur & Rothwell

Late Cretaceous (Campanian)

Horseshoe Canyon Formation

 Canada

A species of Cunninghamia.

Donponoxylon[17]

Gen. et 2 sp. nov.

Valid

Tidwell, Britt & Wright

Middle to Late Jurassic

 Australia
 New Zealand

A seed plant of uncertain phylogenetic placement. The type species is Donponoxylon bennettii; genus also contains Donponoxylon jacksonii.

Dordrechtites arcanus[18]

Sp. nov.

Valid

Bergene, Taylor & Taylor

Middle Triassic (Anisian)

Fremouw Formation

 Antarctica

A gymnosperm of uncertain phylogenetic placement.

Friedsellowia gracilifolia[19]

Gen. nov. et sp. nov.

Valid

Löwe et al.

Early Cretaceous

Crato Formation

 Brazil

A gnetophyte. The type species of a new genus.

Itajuba[20]

Gen. et sp. nov

Valid

Ricardi-Branco et al.

Early Cretaceous (Aptian or Albian)

Crato Member of the Santana Formation

 Brazil

A member of Gnetales. The type species is Itajuba yansanae.

Pagiophyllum heshanense[21]

Sp. nov.

Valid

Liu & Yao

Permian

 China

A species of Pagiophyllum.

Pararaucaria delfueyoi[22]

Sp. nov.

Valid

Escapa et al.

Late Jurassic

Cañadón Calcáreo Formation

 Argentina

A member of Coniferales belonging to the family Cheirolepidiaceae, a species of Pararaucaria.

Prumnopitys anglica[23]

Sp. nov.

Valid

Greenwood, Hill & Conran

Eocene

Branksome Formation

 United Kingdom

A species of Prumnopitys.

Schizolepidopsis canicularis[24]

Sp. nov.

Valid

Leslie et al.

Early Cretaceous (probably Aptian-Albian)

Tevshiin Govi Formation

 Mongolia

A member of Coniferales, possibly a member of the stem group of Pinaceae; a species of Schizolepidopsis.

Stephanospermum braidwoodensis[25]

Sp. nov.

Valid

Spencer, Hilton & Sutton

Carboniferous

Mazon Creek fossil beds

 United States

A member of Medullosales, a species of Stephanospermum.

Stephanospermum shuichengensis[26]

Sp. nov.

Valid

Spencer et al.

Permian (Lopingian)

Wangjiazhai Formation

 China

A member of Medullosales, a species of Stephanospermum.

Other plants

Name Novelty Status Authors Age Unit Location Notes Images

Ashicaulis beipiaoensis[27]

Sp. nov

Valid

Tian et al.

Middle Jurassic

Tiaojishan Formation

 China

A member of Osmundaceae, a species of Ashicaulis.

Bryiidites[28]

Gen. et sp. nov.

Valid

Barclay et al.

Late Cretaceous (late Cenomanian)

Middle Dakota Formation

 United States

A member of Bryidae of uncertain phylogenetic placement. The type species is Bryiidites utahensis.

Franhueberia[29]

Gen. et sp. nov.

Valid

Hoffman & Tomescu

Devonian (Emsian)

Battery Point Formation

 Canada

A basal member of Euphyllophytina. The type species is Franhueberia gerriennei.

Palaeosorum[30]

Gen. et sp. nov

Valid

Jacques & Zhou in Jacques, Su & Zhou

Miocene

Dajie Formation

 China

A member of Polypodiaceae. The type species is Palaeosorum ellipticum.

Yavanna[31]

Gen. et sp. nov

Valid

Vera

Early Cretaceous (Aptian)

Cerro Negro Formation

 Antarctica
(Livingston Island)

A member of Cyatheales of uncertain phylogenetic placement. The type species is Yavanna chimaerica.

Arthropods

Bryozoans

Name Novelty Status Authors Age Unit Location Notes Images

Adlatipora[32]

Gen. et sp. nov

Valid

Gautier, Jackson & McKinney

Permian

 United States

An acanthocladiid bryozoan. The type species is Adlatipora fossulata.

Basyaylella[33]

Gen. et sp. nov

Valid

Zágoršek & Gordon

Late Tortonian

Mut Basin

 Turkey

An ascophoran bryozoan. Its type species is Basyaylella elsae.

Charixa goshouraensis[34]

Sp. nov

Valid[35]

Dick et al.

Cretaceous (late Albian or early Cenomanian)

 Japan

A malacostegan, a species of Charixa.

Conopeum wilsoni[36]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Conopeum.

Haplostoechios[34]

Gen. et 2 sp. nov

Valid[35]

Dick et al.

Cretaceous (late Albian or early Cenomanian)

 Japan

A malacostegan. Genus contains two species: H. hayamiae and H. clusum.

Heloclema uralicum[37]

Sp. nov

Valid

Tolokonnikova

Early Carboniferous

 Russia

A species of Heloclema.

Monoceratopora whybrowi[36]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Monoceratopora.

Ostrovskia[33]

Gen. et sp. nov

Valid

Zágoršek & Gordon

Late Tortonian

Mut Basin

 Turkey

An ascophoran bryozoan. Its type species is Ostrovskia triforamina.

Primorella kodinkensis[37]

Sp. nov

Valid

Tolokonnikova

Early Carboniferous

 Russia

A species of Primorella.

Tyloporella smithi[36]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Tyloporella.

Wilbertopora cheethami[36]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Wilbertopora.

Brachiopods

Name Novelty Status Authors Age Unit Location Notes Images

Arenorthis paranaensis[38]

Sp. nov

Valid

Benedetto, Halpern & Inchausti

Late Ordovician (Hirnantian)

Eusebio Ayala Formation

 Paraguay

A species of Arenorthis.

Askepasma saproconcha[39]

sp nov

Valid

Topper et al..

Lower Cambrian

Wirrapowie Limestone

 Australia

A member of Paterinata, a species of Askepasma.

Biseptata[40]

Gen. et sp. nov

Valid

Madison

Ordovician

 Russia

A member of Strophomenida. The type species is Biseptata briani.

Menathyris[41]

Gen. et sp. nov

Valid

Feldman

Middle Triassic (early Ladinian)

Saharonim Formation

 Israel

A zeilleriid terebratulid. The type species is Menathyris wilsoni.

Neochonetes (Huangichonetes) archboldi[42]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Neochonetes semicircularis[42]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Neochonetes (Sommeriella) rectangularis[42]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Neochonetes (Sommeriella) waterhousei[42]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Plectothyrella? itacurubiensis[38]

Sp. nov

Valid

Benedetto, Halpern & Inchausti

Late Ordovician (Hirnantian)

Eusebio Ayala Formation

 Paraguay

Possibly a species of Plectothyrella.

Proclinorthis[43]

Gen. et sp. nov

Valid

Colmenar, Villas & Vizcaïno

Late Ordovician

Glauzy Formation

 France

A platyorthid. The type species is Proclinorthis vailhanensis.

Pseudocrania insperata[44]

Sp. nov

Valid

Bassett et al.

Middle Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Craniida, a species of Pseudocrania.

Sphenope[45]

Gen. et comb. et sp. nov

Valid

Vörös

Late Jurassic to Early Cretaceous (Tithonian to Berriasian)

 Hungary
 Italy

A pygopid terebratulid. A new genus for "Terebratula" misilmerensis Gemmellaro (1871); genus also contains the new species Sphenope bifida.

Tethyochonetes cheni[42]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Tethyochonetes.

Tethyochonetes sheni[42]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Tethyochonetes.

Molluscs

Echinoderms

Name Novelty Status Authors Age Unit Location Notes Images

Abertella miskellyi[46]

Sp. nov

Valid

Kroh et al.

Miocene

Camarones Formation

 Argentina

An abertellid sand dollar, a species of Abertella.

Alternacantha[47]

Gen. et sp. nov

Valid

Thuy & Meyer

Middle Jurassic

  Switzerland

An ophiacanthid brittle star. The type species is Alternacantha occulta.

Aragocystites[48]

Gen. et sp. nov

Valid

Zamora

Middle Cambrian

Murero Formation

 Spain

An edrioasteroid. The type species is Aragocystites belli.

Bathyovulaster[49]

Gen. et sp. nov

Valid

Smith

Santonian

 Italy

A spatangoid. The type species is Bathyovulaster disjunctus.

Bellastrella[50]

Gen. et sp. nov

Valid

Müller, Hahn and Bohatý

Middle Devonian

 Germany

An agelacrinitid edrioasteroid. The type species is Bellastrella eifeliana.

Ctenocrinus branisai[51]

Sp. nov

Valid

Thompson, Ausich & Smith

Early Devonian (Emsian)

Belèn Formation

 Bolivia

A crinoid, a species of Ctenocrinus.

Cyclolampas altus[52]

Sp. nov

Valid

Saucede et al.

Middle Jurassic (late Callovian)

 France

An echinoid, a species of Cyclolampas.

Eozonella[53]

Gen. et sp. nov

Valid

Thuy, Marty & Comment

Late Jurassic (late Oxfordian)

  Switzerland

An ophiolepidid brittle star. The type species is Eozonella bergeri.

Graciacystis[54]

Gen. et sp. nov

Valid

Zamora, Rahman & Smith

Middle Cambrian

 Spain

A basal cinctan echinoderm. The type species is Graciacystis ambigua.

Griphocrinus pirovanoi[51]

Sp. nov

Valid

Thompson, Ausich & Smith

Early Devonian (Emsian)

Icla Formation

 Bolivia

A crinoid, a species of Griphocrinus.

Hanshessia[47]

Gen. et sp. nov

Valid

Thuy & Meyer

Middle Jurassic

  Switzerland

An ophiacanthid brittle star. The type species is Hanshessia trochitophila.

Juracantha[53]

Gen. et sp. nov

Valid

Thuy, Marty & Comment

Late Jurassic (late Oxfordian)

  Switzerland

An ophiacanthid brittle star. The type species is Juracantha hottingeri.

Kosachenkoastrus[55]

Gen. et sp. nov

Valid

Rozhnov

Middle Ordovician

 Russia

A member of Parablastoidea. The type species is Kosachenkoastrus volkhovensis.

Lutocrinus[51]

Gen. et sp. nov

Valid

Thompson, Ausich & Smith

Early Devonian (Emsian)

Icla Formation

 Bolivia

A crinoid. The type species is Lutocrinus boliviaensis.

Ophiotholia aurora[47]

Sp. nov

Valid

Thuy & Meyer

Middle Jurassic

  Switzerland

An ophiomycetid brittle star, a species of Ophiotholia.

Rhenopyrgus piojoensis[56]

Sp nov

Valid

Sumrall, Heredia, Rodríguez & Mestre

Silurian (lower Ludlow)

Los Espejos Formation

 Argentina

A rhenopyrgid edrioasteroid, a species of Rhenopyrgus.

Simakocrinus[57]

Gen. et 2 sp. nov.

Valid

Prokop

Devonian (Pragian, Emsian and Eifelian)

 Czech Republic

A crinoid belonging to the group Disparida, possibly a member of the family Myelodactylidae. The type species is Simakocrinus facilis sp. nov (col.); genus also contains Simakocrinus diligens sp. nov (col.).

Conodonts

Name Novelty Status Authors Age Unit Location Notes Images

Acanthodus humachensis[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Acanthodus.

Acanthodus raqueli[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Acanthodus.

Acodus primitivus[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Acodus.

Acuminatella[59]

Gen. et 2 sp. nov

Valid

Orchard

Late Triassic (late Carnian to early Norian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. The type species is Acuminatella acuminata; genus also contains Acuminatella angusta.

Bispathodus ultimus bartzschi[60]

Subsp nov

Valid

Kononova & Weyer

Devonian (late Famennian)

 Germany

A subspecies of Bispathodus ultimus.

Carnepigondolella eozoae[59]

Sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Ludington Formation

 Canada

A gondolellid ozarkodinid, a species of Carnepigondolella.

Carnepigondolella medioconstricta[59]

Sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Ludington Formation

 Canada

A gondolellid ozarkodinid, a species of Carnepigondolella.

Furnishina leei[61]

Sp. nov

Valid

Lee

Cambrian

Machari Formation

 South Korea

Histiodella wuhaiensis[62]

Sp. nov

Valid

Wang et al.

Ordovician

Lower Klimoli Formation

 China

Idiognathodus craticulinodosus[63]

Sp. nov

Valid

Ishida, Suzuki & Inada

Carboniferous (middle–late Bashkirian to early Moscovian)

Ko-yama Limestone Group

 Japan

Kallidontus gondwanicus[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Kallidontus.

Kraussodontus[59]

Gen. et sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. The type species is Kraussodontus peteri.

Mesogondolella daheshenensis[64]

Sp nov

Valid

Wang in Zhou et al.

Permian (probably Wordian)

Daheshen Formation

 China

A species of Mesogondolella.

Mesogondolella subgracilis[64]

Sp nov

Valid

Wang in Zhou et al.

Permian (probably Wordian)

Daheshen Formation

 China

A species of Mesogondolella.

Muellerilepis[65]

Nom. nov

Valid

Bardashev & Bardasheva

Middle Devonian (Givetian)

 Tajikistan

A replacement name for Muellerina Bardashev & Bardasheva (2012) (preoccupied).

Naimanodus[66]

Gen. et sp. nov

Valid

Tolmacheva

Middle Ordovician

 Kazakhstan
 Kyrgyzstan
 Russia

The type species is Naimanodus degtyarevi.

Neopolygnathus margaretae[60]

Sp nov

Valid

Kononova & Weyer

Devonian (late Famennian)

 Germany

A species of Neopolygnathus.

Notiodella[67]

Gen. et sp. nov

Valid

Aldridge et al.

Late Ordovician

Soom Shale Member, Cedarberg Formation

 South Africa

A member of Prioniodontida belonging to the family Balognathidae. The type species is Notiodella keblon.

Palmatolepis khaensis[68]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Palmatolepis marki[68]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Palmatolepis subperlobata lapoensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Parapetella[59]

Gen. et sp. nov

Valid

Orchard

Late Triassic (late Carnian to early Norian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. The type species is Parapetella prominens.

Polygnathus burretti[68]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus communis hanensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus communis namdipensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus communis phaphaensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus crassulus salapensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus extralobatus phoensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus hojedki[69]

Sp. nov

Valid

Gholamalian et al.

Devonian

 Iran

A species of Polygnathus.

Polygnathus pseudocostatus[70]

Sp nov

Valid

Klapper & Vodrážková

Devonian

 United States

A species of Polygnathus.

Primatella[59]

Gen. et comb. et 2 sp. nov

Valid

Orchard

Late Triassic (late Carnian to early Norian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. A new genus for "Epigondolella" primitia Mosher (1970); genus also contains new species Primatella asymmetrica and Primatella conservativa.

Pseudopolygnathus granulosus laepensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Pseudopolygnathus granulosus salawinensis[68]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Pseudopolygnathus maepoensis[68]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Quadralella[59]

Gen. et sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Ludington Formation

 Canada

A gondolellid ozarkodinid. The type species is Quadralella lobata.

Siphonodella banraiensis[68]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Tilcarodus[58]

Gen. et comb. nov

Valid

Zeballo & Albanesi

Early Ordovician

Rupasca Formation

 Argentina

A new genus for "Utahconus" humahuacensis Albanesi and Aceñolaza (2005).

Trapezognathus? primitivus[71]

Sp. nov

Valid

Voldman, Albanesi & Zeballo in Voldman et al.

Early Ordovician

Acoite Formation

 Argentina

Originally described as a possible species of Trapezognathus; subsequently made the type species of a separate genus Zentagnathus.[72]

Utahconus purmamarcensis[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Utahconus.

Utahconus scandodiformis[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Utahconus.

Utahconus tortibasis[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Utahconus.

Variabiloconus crassus[58]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Variabiloconus.

Fishes

Amphibians

Newly named temnospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Broiliellus reiszi[73]

Sp. nov

Valid

Holmes, Berman & Anderson

Early Permian

El Cobre Canyon Formation

 United States

A dissorophid, a species of Broiliellus.

Parotosuchus ptaszynskii[74]

Sp nov

Valid

Sulej & Niedźwiedzki

Late Olenekian

 Poland

A species of Parotosuchus.

Reiszerpeton[75]

Gen. et sp. nov

Valid

Maddin et al.

Early Permian

Archer City Formation

 United States

A dissorophid. The type species is Reiszerpeton renascentis.

Scapanops[76]

Gen. et sp. nov

Valid

Schoch & Sues

Early Permian

Nocona Formation

 United States

A dissorophid. The type species is Scapanops neglecta.

Tersomius dolesensis[77]

Sp nov

Valid

Anderson & Bolt

Early Permian

 United States

A species of Tersomius.

Trimerorhachis greggi[78]

Sp nov

Valid

Milner & Schoch

Early Permian

 United States

A species of Trimerorhachis.

Newly named lepospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Batropetes niederkirchensis[79]

Sp. nov

Valid

Glienke

Early Permian (Rotliegend)

Saar–Nahe Basin

 Germany

A species of Batropetes.

Huskerpeton[80]

Gen. et sp. nov

Valid

Huttenlocker et al.

Early Permian

Eskridge Formation

 United States

A recumbirostran. The type species is Huskerpeton englehorni.

Proxilodon[80]

Gen. et comb. nov

Valid

Huttenlocker et al.

Early Permian

Speiser Formation

 United States

A recumbirostran, a new genus for "Euryodus" bonneri.

Newly named lissamphibians

Name Novelty Status Authors Age Unit Location Notes Images

Aerugoamnis[81]

Gen. et sp. nov

Valid

Henrici, Báez & Grande

Early Eocene (Wasatchian)

Green River Formation

 United States

A relative of parsley frogs. The type species is Aerugoamnis paulus.

Eobarbourula[82]

Gen. et sp. nov

Valid

Folie et al..

Early Eocene

Cambay Shale Formation

 India

A bombinatorid. Its type species is Eobarbourula delfinoi.

Gracilibatrachus[83]

Gen. et sp. nov

Valid

Báez

Early Cretaceous (late Barremian)

 Spain

A relative of the family Pipidae. The type species is Gracilibatrachus avallei.

Iberobatrachus[83]

Gen. et sp. nov

Valid

Báez

Early Cretaceous (late Barremian)

 Spain

A close relative of the genus Discoglossus. The type species is Iberobatrachus angelae.

Ichthyosaura randeckensis[84]

Sp. nov

Valid

Schoch & Rasser

Miocene

 Germany

A newt related to the Alpine newt.

Indorana[82]

Gen. et sp. nov

Valid

Folie et al..

Early Eocene

Cambay Shale Formation

 India

A possible rhacophorid. Its type species is Indorana prasadi.

Leiopelma acricarina[85]

Sp. nov

Valid

Worthy et al.

Early Miocene

 New Zealand

A member of Leiopelmatidae.

Leiopelma miocaenale[85]

Sp. nov

Valid

Worthy et al.

Early Miocene

 New Zealand

A member of Leiopelmatidae.

Liaobatrachus zhaoi[86]

Sp. nov

Valid

Dong et al..

Early Cretaceous (Barremian)

Yixian Formation

 China

An anuran of uncertain phylogenetic placement, a species of Liaobatrachus.

Paranecturus[87]

Gen. et sp. nov

Valid

Demar

Late Cretaceous (Maastrichtian)

Hell Creek Formation

 United States

A mudpuppy. The type species is Paranecturus garbanii.

Ukrainurus[88]

Gen. et sp. nov

Valid

Vasilyan et al.

Miocene

 Ukraine

A relative of cryptobranchids. The type species is Ukrainurus hypsognathus.

Wesserpeton[89]

Gen. et sp. nov

Valid

Sweetman & Gardner

Barremian

Wessex Formation

 England

An albanerpetontid.

Turtles

Newly named turtles

Name Novelty Status Authors Age Unit Location Notes Images

Adocus inexpectatus[90]

Sp. nov

Valid

Danilov et al.

Late Eocene

Youganwo Formation

 China

An adocid, a species of Adocus.

Atoposemys[91]

Gen. et sp. nov

Valid

Hutchison

Early Paleocene (Puercan)

Fort Union Formation

 United States

A trionychid. The type species is Atoposemys entopteros.

Bairdemys healeyorum[92]

Sp. nov

Valid

Weems & Knight

Late Oligocene

Chandler Bridge Formation

 United States

A podocnemidid pleurodiran, a species of Bairdemys.

Basilemys gaffneyi[93]

Sp. nov

Valid

Sullivan, Jasinski & Lucas

Late Campanian

 United States

A nanhsiungchelyid turtle, a species of Basilemys.

Brachyopsemys[94]

Gen. et sp. nov

Valid

Tong & Meylan

Early Paleocene

Ouled Abdoun Basin

 Morocco

A cryptodiran closely related to Sandownia. The type species is Brachyopsemys tingitana.

Camerochelys[95]

Gen. et sp. nov

Valid

Pérez-García & Murelaga

Early Cretaceous (Hauterivian or Barremian)

Cameros Basin

 Spain

A member of Pan-Cryptodira (the clade containing living cryptodirans and all turtles sharing a more recent common ancestor with them than with pleurodirans), possibly a xinjiangchelyid. The type species is Camerochelys vilanovai.

Cardichelyon[91]

Gen. et sp. nov

Valid

Hutchison

Early Eocene (Wasatchian)

 United States

A platysternid. The type species is Cardichelyon rogerwoodi.

?Centrochelys marocana[96]

Sp. nov

Valid

Gmira et al.

Pliocene

 Morocco

A tortoise, possibly a species of Centrochelys.

Changmachelys[97]

Gen. et sp. nov

Valid

Brinkman et al.

Early Cretaceous

Xiagou Formation

 China

A member of (likely paraphyletic) group "Macrobaenidae". The type species is Changmachelys bohlini.

Chelodina (Chelodina) murrayi[98]

Sp. nov

Valid

Yates

Late Miocene

Waite Formation

 Australia

A chelid, a species of Chelodina.

Chelonoidis lutzae[99]

Sp. nov

Valid

Zacarías et al.

Late Pleistocene (58–22 ka)

 Argentina

A tortoise, a species of Chelonoidis.

Chrysemys isoni[100]

Sp. nov

Valid

Weems & George

Miocene

Calvert Formation

 United States

A relative of the painted turtle.

Cuora chiangmuanensis[101]

Sp. nov

Valid

Naksri et al.

Late Middle or early Late Miocene

 Thailand

An Asian box turtle.

Floridemys hurdi[100]

Sp. nov

Valid

Weems & George

Miocene

Calvert Formation

 United States

A tortoise, a species of Floridemys.

Kappachelys[102]

Gen. et sp. nov

Valid

Hirayama, Isaji & Hibino

Early Cretaceous

Akaiwa Formation

 Japan

A relative of trionychids. The type species is Kappachelys okurai.

Laganemys[103]

Gen. et sp. nov

Valid

Sereno & ElShafie

Aptian or Albian

Elrhaz Formation

 Niger

An araripemydid pleurodiran. The type species is Laganemys tenerensis.

Neochelys liriae[104]

Sp. nov

Valid

Pérez-García & de Lapparent de Broin

Eocene (Ypresian)

 France

A podocnemidid, a species of Neochelys.

Neurankylus lithographicus[105]

Sp. nov

Valid

Larson et al.

Santonian

Milk River Formation

 Canada

A neurankyline baenid, a species of Neurankylus.

Nostimochelone[106]

Gen. et sp. nov

Valid

Georgalis et al.

Early Miocene

Zeugostasion Formation

 Greece

A pleurodiran, a member of Podocnemidoidea. The type species is Nostimochelone lampra.

Ocadia tanegashimensis[107]

Sp. nov

Valid

Takahashi et al.

Early middle Miocene

Kawachi Formation

 Japan

A geoemydid, a species of Ocadia.

Ocepechelon[108]

Gen. et sp. nov

Valid

Bardet et al.

Late Cretaceous (late Maastrichtian)

Oulad Abdoun Basin

 Morocco

A relative of dermochelyids and protostegids. The type species is Ocepechelon bouyai.

Paramongolemys[109]

Gen. et sp. nov

Valid

Danilov & Sukhanov

Late Paleocene

Naranbulak Formation

 Mongolia

A basal member of Testudinoidea. The type species is Paramongolemys khosatzkyi.

Planetochelys dithyros[91]

Sp. nov

Valid

Hutchison

Early Eocene (Wasatchian)

 United States

A relative of trionychids, a species of Planetochelys.

Psilosemys[91]

Gen. et sp. nov

Valid

Hutchison

Early Eocene (Wasatchian)

 United States

The earliest North American emydid. The type species is Psilosemys wyomingensis.

Scabremys[93]

Gen. et comb. nov

Valid

Sullivan, Jasinski & Lucas

Late Campanian

 United States

A baenid cryptodiran, a new genus for "Baena" ornata Gilmore (1935).

Shandongemys[110]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous

 China

A lindholmemydid testudinoid. The type species is Shandongemys dongwuica.

Sinemys brevispinus[111]

Sp. nov

Valid

Tong & Brinkman

Early Cretaceous

Luohandong Formation

 China

A sinemydid cryptodiran, a species of Sinemys.

Spoochelys[112]

Gen. et sp. nov

Valid

Smith & Kear

Early or middle Albian

Griman Creek Formation

 Australia

A meiolaniid-like turtle. The type species is Spoochelys ormondea.

Testudo oughlamensis[96]

Sp. nov

Valid

Gmira et al.

Pliocene

 Morocco

A tortoise, a species of Testudo.

Trapalcochelys[113]

Gen. et sp. nov

Valid

Sterli, de la Fuente & Cerda

Late Cretaceous (late Campanian to early Maastrichtian)

Allen Formation

 Argentina

A relative of meiolaniids. The type species is Trapalcochelys sulcata.

Tullochelys[91]

Gen. et sp. nov

Valid

Hutchison

Maastrichtian to early Paleocene (Puercan)

Fort Union Formation
Hell Creek Formation

 United States

A chelydrid. The type species is Tullochelys montanus.

Xenochelys floridensis[114]

Sp. nov

Valid

Bourque

Late Oligocene (early Arikareean)

 United States

A kinosternid, a species of Xenochelys.

Xinjiangchelys radiplicatoides[115]

Sp. nov

Valid

Brinkman et al.

Late Middle or early Late Jurassic

Shishugou Formation

 China

A species of Xinjiangchelys.

Xinjiangchelys wusu[116]

Sp. nov

Valid

Rabi et al.

Middle Jurassic

?Qigu Formation

 China

A species of Xinjiangchelys.

Thalattosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Concavispina[117]

Gen. et sp. nov

Valid

Zhao et al.

Triassic

Xiaowa Formation

 China

A thalattosaurid thalattosaur. The type species is Concavispina biseridens

Ichthyopterygians

Name Novelty Status Authors Age Unit Location Notes Images

Chaohusaurus zhangjiawanensis[118]

Sp. nov

Valid

Chen et al.

Early Triassic

Jialingjiang Formation

 China

A species of Chaohusaurus.

Gulosaurus[119]

Gen. et sp. nov

Valid

Cuthbertson, Russell & Anderson

Early Triassic

Sulphur Mountain Formation

 Canada

A grippidian. The type species is Gulosaurus helmi.

Malawania[120]

Gen. et sp. nov

Valid

Fischer et al.

Early Cretaceous (late Hauterivian to Barremian)

 Iraq

A basal member of Thunnosauria. The type species is Malawania anachronus.

Thalattoarchon[121]

Gen. et sp. nov

Valid

Fröbisch et al.

Middle Triassic

 United States

A merriamosaur ichthyosaur. The type species is Thalattoarchon saurophagis

Lepidosauromorphs

Newly named sauropterygians

Name Novelty Status Authors Age Unit Location Notes Images

Cryonectes[122]

Gen. et sp. nov

Valid

Vincent, Bardet & Mattioli

Pliensbachian

 France

A pliosaurid. The type species is Cryonectes neustriacus

Gronausaurus[123]

Gen. et sp. nov

Valid

Hampe

Early Cretaceous (Berriasian)

 Germany

A leptocleidid[123] or an elasmosaurid.[124] The type species is Gronausaurus wegneri

Megacephalosaurus[125]

Gen. et sp. nov

Valid

Schumacher, Carpenter & Everhart

Late Cretaceous (middle Turonian)

 United States

A pliosaurid. The type species is Megacephalosaurus eulerti.

Palatodonta[126]

Gen. et sp. nov

Valid

Neenan, Klein & Scheyer

Early Middle Triassic

 Netherlands

A relative of placodonts. The type species is Palatodonta bleekeri.

Pliosaurus carpenteri[124]

Sp. nov

Valid

Benson et al.

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A pliosaurid, a species of Pliosaurus

Pliosaurus kevani[124]

Sp. nov

Valid

Benson et al.

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A pliosaurid, a species of Pliosaurus

Pliosaurus westburyensis[124]

Sp. nov

Valid

Benson et al.

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A pliosaurid, a species of Pliosaurus

Serpianosaurus germanicus[127]

Sp. nov

Valid

Diedrich

Middle Triassic (Anisian)

 Germany

A pachypleurosaur, a species of Serpianosaurus.

Newly named rhynchocephalians

Name Novelty Status Authors Age Unit Location Notes Images

Sphenotitan[128]

Gen. et sp. nov

Valid

Martínez et al.

Late Triassic

Quebrada del Barro Formation

 Argentina

An opisthodontian rhynchocephalian. The type species is Sphenotitan leyesi.

Newly named lizards

Name Novelty Status Authors Age Unit Location Notes Images

Apsgnathus[129]

Gen. et sp. nov

Valid

Nydam, Rowe & Cifelli

Late Cretaceous (late Campanian, Judithian)

Aguja Formation

 United States

A member of Scincomorpha of uncertain phylogenetic placement. The type species is Apsgnathus triptodon.

Arcanosaurus[130]

Gen. et sp. nov

Valid

Houssaye et al.

Early Cretaceous (Barremian or Aptian)

Cameros Basin

 Spain

An anguimorph lizard, possibly a varanoid. The type species is Arcanosaurus ibericus.

Arpadosaurus sepulchralis[131]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Glyptosaurinae, a species of Arpadosaurus.

Barbaturex[132]

Gen. et sp. nov

Valid

Head et al.

Late middle Eocene (37.2 ± 1.3 Ma)

Pondaung Formation

 Myanmar

An iguanian lizard related to the genus Uromastyx. The type species is Barbaturex morrisoni.

Bicuspidon smikros[133]

Sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A member of Polyglyphanodontinae, a species of Bicuspidon.

Blanosaurus[134]

Gen. et sp. nov

Valid

Folie, Smith & Smith

Early Eocene

 Belgium
 France

An amphisbaenian. The type species is Blanosaurus primeocaenus.

Camptognathosaurus[134]

Gen. et sp. nov

Valid

Folie, Smith & Smith

Late Paleocene

 France

An amphisbaenian. The type species is Camptognathosaurus parisiensis.

Catactegenys[129]

Gen. et sp. nov

Valid

Nydam, Rowe & Cifelli

Late Cretaceous (late Campanian, Judithian)

Aguja Formation

 United States

A night lizard. The type species is Catactegenys solaster.

Cemeterius[135][136]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A platynotan lizard of uncertain phylogenetic placement. The type species is Cemeterius monstrosus.

Cnodontosaurus[133]

Gen. et sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A member of Anguimorpha related to xenosaurids. The type species is Cnodontosaurus suchockii.

Dakotasaurus[133]

Gen. et sp. nov

Junior homonym

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.[133] The type species is Dakotasaurus gillettorum. The generic name turned out to be preoccupied by Dakotasaurus Branson & Mehl (1932); the lizard genus was subsequently renamed Dakotaseps.[137]

Desertiguana[138]

Gen. et sp. nov

Valid

Alifanov

Late Cretaceous

 Mongolia

A phrynosomatid lizard. The type species is Desertiguana gobiensis.

Distortodon[139]

Gen. et sp. nov

Valid

Makádi

Late Cretaceous (Santonian)

 Hungary

A member of Polyglyphanodontinae. The type species is Distortodon rhomboideus.

Entomophontes[131]

Gen. et 2 sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A relative of species belonging to the genus Xenosaurus. Genus contains two species: Entomophontes incrustatus and Entomophontes hutchisoni.

Eonatator coellensis[140]

Sp. nov

Valid

Páramo

Campanian

 Colombia

A mosasaur, a new species of Eonatator.

Glyptosaurus agmodon[131]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Glyptosaurinae, a species of Glyptosaurus.

Heterodontagama[141]

Gen. et sp. nov

Valid

Rana et al.

Early Eocene (early to middle Ypresian)

Cambay Shale Formation

 India

A member of Priscagamidae. The type species is Heterodontagama borsukae.

Indiagama[141]

Gen. et sp. nov

Valid

Rana et al.

Early Eocene (early to middle Ypresian)

Cambay Shale Formation

 India

A member of Agamidae. The type species is Indiagama gujarata.

Lamiasaura[135][136]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A squamate of uncertain phylogenetic placement. The type species is Lamiasaura ferox.

Lonchisaurus[135][136]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A scincomorph lizard of uncertain phylogenetic placement. The type species is Lonchisaurus trichurus.

Monocnemodon[133]

Gen. et sp. nov

Valid

Nydam

Late Cretaceous (Santonian)

Straight Cliffs Formation

 United States

A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.[133] The type species is Monocnemodon syphakos.

Obamadon[135][136]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Hell Creek Formation
Lance Formation

 United States

A polyglyphanodontian lizard of uncertain phylogenetic placement. The type species is Obamadon gracilis.

Odaxosaurus roosevelti[133]

Sp. nov

Valid

Nydam

Late Cretaceous

Kaiparowits Formation

 United States

A member of Anguidae, a species of Odaxosaurus.

Palaeoxantusia amara[131]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A night lizard, a species of Palaeoxantusia.

Palepidophyma[131]

Gen. et 2 sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A night lizard. Genus contains two species: Palepidophyma paradisa and Palepidophyma lilliputiana.

Parasaniwa cynochoros[133]

Sp. nov

Valid

Nydam

Late Cretaceous

Kaiparowits Formation

 United States

A member of Platynota of uncertain phylogenetic placement, a species of Parasaniwa.

Pariguana[135][136]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

An iguanid lizard of uncertain phylogenetic placement. The type species is Pariguana lancensis.

Pelsochamops[142]

Gen. et sp. nov

Valid

Makádi

Late Cretaceous (Santonian)

Csehbánya Formation

 Hungary

A chamopsiid. The type species is Pelsochamops infrequens.

Plesiolacerta eratosthenesi[143]

Sp. nov

Valid

Čerňanský & Augé

Late Oligocene

 Germany

A lacertid lizard, a species of Plesiolacerta.

Polyodontobaena[134]

Gen. et sp. nov

Valid

Folie, Smith & Smith

Early Paleocene

Hainin Formation

 Belgium

An amphisbaenian. The type species is Polyodontobaena belgica.

Provaranosaurus fatuus[131]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A relative of the Chinese crocodile lizard, a species of Provaranosaurus.

Romeosaurus[144]

Gen. et 2 sp. nov

Valid

Palci, Caldwell & Papazzoni

Late Cretaceous (early Turonian to early Santonian)

 Italy

A mosasaur related to Russellosaurus and Yaguarasaurus. Genus contains two species: Romeosaurus fumanensis and Romeosaurus sorbinii.

Schillerosaurus[145]

Nom. nov

Valid

Nydam, Chure & Evans

Late Jurassic

Morrison Formation

 United States

A replacement name for Schilleria Evans & Chure (1999).

Scincoideus grassator[131]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Lacertiformes, a species of Scincoideus.

Socognathus brachyodon[135][136]

Sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A chamopsiid polyglyphanodontian lizard, a species of Socognathus.

Suratagama[141]

Gen. et sp. nov

Valid

Rana et al.

Early Eocene (early to middle Ypresian)

Cambay Shale Formation

 India

A member of Agamidae. The type species is Suratagama neeraae.

Suzanniwana revenanta[131]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Pleurodonta, a species of Suzanniwana.

Tiliqua laticephala[146]

Sp. nov

Valid

Čerňanský & Hutchinson

Pliocene

 Australia

A blue-tongued skink.

Utahgenys antongai[133]

Sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A contogeniid scincomorph, a species of Utahgenys.

Webbsaurus[133]

Gen. et sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A scincomorph related to skinks. The type species is Webbsaurus lofgreni.

Xestops savagei[131]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Glyptosaurinae, a species of Xestops.

Newly named snakes

Name Novelty Status Authors Age Unit Location Notes Images

Cerberophis[135][136]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Hell Creek Formation

 United States

An alethinophidian snake of uncertain phylogenetic placement. The type species is Cerberophis robustus.

Kataria[147]

Gen. et sp. nov

Valid

Scanferla et al.

Early Paleocene (Danian)

Santa Lucía Formation

 Bolivia

A macrostomatan snake related to caenophidians. The type species is Kataria anisodonta.

Nidophis[148]

Gen. et sp. nov

Valid

Vasile, Csiki-Sava & Venczel

Late Cretaceous (Maastrichtian)

Haţeg Basin

 Romania

A madtsoiid. The type species is Nidophis insularis.

Archosauromorphs

Newly named basal archosauromorphs

Name Novelty Status Authors Age Unit Location Notes Images

Ankylosuchus[149]

Gen. et sp. nov

Valid

Lucas, Spielmann & Hunt

Late Triassic

Colorado City Formation

 United States

A doswelliid. The type species is Ankylosuchus chinlegroupensis.

Asperoris[150]

Gen. et sp. nov

Valid

Nesbitt, Butler & Gower

Middle Triassic (probably late Anisian)

Manda beds

 Tanzania

A non-archosaurian archosauriform of uncertain phylogenetic placement. The type species is Asperoris mnyama.

Fuyuansaurus[151]

Gen. et sp. nov

Valid

Fraser, Rieppel & Chun

Middle Triassic (Ladinian)

Falang Formation

 China

A protorosaur. The type species is Fuyuansaurus acutirostris.

Jaxtasuchus[152]

Gen. et sp. nov

Valid[153]

Schoch & Sues

Middle Triassic (Ladinian)

Erfurt Formation

 Germany

A doswelliid. The type species is Jaxtasuchus salomoni.

Machaeroprosopus lottorum[154]

Sp. nov

Valid

Hungerbühler et al.

Late Triassic

Cooper Canyon Formation

 United States

A phytosaur, a species of Machaeroprosopus.

Macrocnemus obristi[155]

Sp. nov

Valid

Fraser & Furrer

Middle Triassic (Ladinian)

Prosanto Formation

  Switzerland

A protorosaur, a species of Macrocnemus.

Wannia[156]

Gen. et comb. nov

Valid

Stocker

Late Triassic

 United States

A phytosaur; a new genus for "Paleorhinus" scurriensis Langston (1949).

Archosaurs

Other reptiles

Name Novelty Status Authors Age Unit Location Notes Images

Dolerosaurus[157]

Gen. et comb. nov

Valid

Butler

Late Triassic (late Carnian)

Lunz Formation

 Austria

A diapsid of uncertain phylogenetic placement, a new genus for "Francosuchus" trauthi Huene (1939).

Ruhuhuaria[158]

Gen. et sp. nov

Valid

Tsuji, Sobral & Müller

Middle Triassic

Manda Formation

 Tanzania

A procolophonoid parareptile, possibly an owenettid. The type species is Ruhuhuaria reiszi.

Synapsids

Non-mammalian synapsids

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Diegocanis[160]

Gen. et sp. nov

Valid

Martínez, Fernandez & Alcober

Late Triassic

Ischigualasto Formation

 Argentina

A member of Eucynodontia closely related to Ecteninion and Trucidocynodon. The type species is Diegocanis elegans.

Mandagomphodon[161]

Gen. et comb. nov

Valid[162]

Hopson

Middle Triassic

Manda Formation

 Tanzania

A traversodontid cynodont, a new genus for "Scalenodon" hirschsoni Crompton (1972).

Niassodon[163]

Gen. et sp. nov

Valid

Castanhinha et al.

Late Permian

 Mozambique

A kingoriid emydopoid dicynodont. The type species is Niassodon mfumukasi.

Tambacarnifex[164]

Gen. et sp. nov

Valid[162]

Berman et al.

Early Permian

Tambach Formation

 Germany

A varanodontine varanopid. The type species is Tambacarnifex unguifalcatus.

Mammals

Other animals

Name Novelty Status Authors Age Unit Location Notes Images

Anomalocaris kunmingensis[165]

Sp. nov

Valid

Wang, Huang & Hu

Cambrian

Wulongqing Formation

 China

A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods), a species of Anomalocaris.

Aulastraea crassicalix[166]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Aulastraea.

Cambrocornulitus[167]

Gen. et sp. nov

Valid

Yang et al.

Early Cambrian

Qiongzhusi Formation

 China

An animal of uncertain phylogenetic placement, possibly a member of Cornulitida. The type species is Cambrocornulitus rarus.

Conularia porcella[168]

Sp. nov

Valid

Robson & Young

Late Ordovician (middle Katian)

Red River Formation

 Canada

A conulariid, a species of Conularia.

Gracilitheca astronauta[169]

Sp. nov

Valid

Valent & Fatka

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.

Gracilitheca mirabilis[170]

Sp. nov

Valid

Valent, Fatka & Marek

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.

Gracilitheca triangularis[170]

Sp. nov

Valid

Valent, Fatka & Marek

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha, a species of Gracilitheca.

Heteropistophyllum[166]

Gen. et comb. nov

Valid

Löser, Castro & Nieto

Early Cretaceous

 Slovenia
 Spain

A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. A new genus for "Pseudopistophyllum" quinqueseptatum Turnšek & Buser (1976).

Metaulastraea elae[166]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Metaulastraea.

Microconchus utahensis[171]

Sp. nov

Valid

Zatoń, Taylor & Vinn

Early Triassic

Virgin Formation

 United States

A member of Microconchida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Microconchus.

Mitrodendron ultimus[166]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Mitrodendron.

Nephrotheca betula[170]

Sp. nov

Valid

Valent, Fatka & Marek

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha, a species of Nephrotheca.

Nephrotheca sophia[169]

Sp. nov

Valid

Valent & Fatka

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha, a species of Nephrotheca.

Paraacanthogyra hispanica[166]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Paraacanthogyra.

Paraconularia bachmanni[172]

Sp. nov

Valid

Barth et al.

Late Triassic

 Germany

A conulariid, a species of Paraconularia.

Paranomalocaris[165]

Gen. et sp. nov

Valid

Wang, Huang & Hu

Cambrian

Wulongqing Formation

 China

A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods). The type species is Paranomalocaris multisegmentalis.

Pleurodendron[166]

Gen. et sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. The type species is Pleurodendron prebeticum.

Quasiaulacera[173]

Gen. et 2 sp. nov

Valid

Copper, Stock & Jin

Late Ordovician (Hirnantian)

Ellis Bay Formation

 Canada

A stromatoporoid. Genus contains two species: Quasiaulacera stellata and Q. occidua.

Silingastraea valentina[166]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Silingastraea.

Spinitheca[174]

Gen. et sp. nov

Valid

Novozhilova

Early Cambrian (Atdabanian)

 Russia

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs). The type species is Spinitheca sysoievi.

Other organisms

Name Novelty Status Authors Age Unit Location Notes Images

Honeggeriella[175]

Gen. et sp. nov

Valid

Matsunaga, Stockey & Tomescu

Early Cretaceous (Valanginian-Hauterivian boundary)

Longarm Formation

 Canada

A heteromerous lichen. The type species is Honeggeriella complexa.

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  132. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  138. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  139. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  143. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  144. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  145. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  146. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  147. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  148. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  151. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  152. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  155. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  156. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  157. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  158. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  161. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  164. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  168. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  169. 169.0 169.1 Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
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  174. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).
  175. Lua error in Module:Citation/CS1/Identifiers at line 47: attempt to index field 'wikibase' (a nil value).