Neolithic Europe refers to a prehistoric period in which Neolithic technology was present in Europe. This corresponds roughly to a time between 7000 BCE (the approximate time of the first farming societies in Greece) and c. 1700 BCE (the beginning of the Bronze Age in northwest Europe). The Neolithic overlaps the Mesolithic and Bronze Age periods in Europe as cultural changes moved from the southeast to northwest at about 1 km/year - this is called Neolithic Expansion.
The duration of the Neolithic varies from place to place, its end marked by the introduction of bronze implements: in southeast Europe it is approximately 4,000 years (i.e. 7000 BCE–3000 BCE) while in Northwest Europe it is just under 3,000 years (c. 4500 BCE–1700 BCE).
Basic cultural characteristics
Regardless of specific chronology, many European Neolithic groups share basic characteristics, such as living in small-scale, family-based communities, subsisting on domesticated plants and animals supplemented with the collection of wild plant foods and with hunting, and producing hand-made pottery, that is, pottery made without the potter's wheel. Polished stone axes lie at the heart of the neolithic (new stone) culture, enabling forest clearance for agriculture and production of wood for dwellings, as well as fuel.
There are also many differences, with some Neolithic communities in southeastern Europe living in heavily fortified settlements of 3,000-4,000 people (e.g., Sesklo in Greece) whereas Neolithic groups in Britain were small (possibly 50-100 people) and highly mobile cattle-herders.
The details of the origin, chronology, social organization, subsistence practices and ideology of the peoples of Neolithic Europe are obtained from archaeology, and not historical records, since these people left none. Since the 1970s, population genetics has provided independent data on the population history of Neolithic Europe, including migration events and genetic relationships with peoples in South Asia.
A further independent tool, linguistics, has contributed hypothetical reconstructions of early European languages and family trees with estimates of dating of splits, in particular theories on the relationship between speakers of Indo-European languages and Neolithic peoples. Some archaeologists believe that the expansion of Neolithic peoples from southwest Asia into Europe, marking the eclipse of Mesolithic culture, coincided with the introduction of Indo-European speakers, whereas other archaeologists and many linguists believe the Indo-European languages were introduced from the Pontic-Caspian steppe during the succeeding Bronze Age.
Archeologists believe that food-producing societies first emerged in the Levantine region of southwest Asia at the close of the mini-Ice Age around 12,000 BCE, and developed into a number of regionally distinctive cultures by the eighth millennium BCE. Remains of food-producing societies in the Aegean have been carbon-dated to around 6500 BCE at Knossos, Franchthi Cave, and a number of mainland sites in Thessaly. Neolithic groups appear soon afterwards in the Balkans and south-central Europe. The Neolithic cultures of southeastern Europe (the Balkans and the Aegean) show some continuity with groups in southwest Asia and Anatolia (e.g., Çatalhöyük).
Current evidence suggests that Neolithic material culture was introduced to Europe via western Anatolia, and that similarities in cultures of North Africa and the Pontic steppes are due to diffusion out of Europe. All Neolithic sites in Europe contain ceramics, and contain the plants and animals domesticated in Southwest Asia: einkorn, emmer, barley, lentils, pigs, goats, sheep, and cattle. Genetic data suggest that no independent domestication of animals took place in Neolithic Europe, and that all domesticated animals were originally domesticated in Southwest Asia. The only domesticate not from Southwest Asia was broomcorn millet, domesticated in East Asia. The earliest evidence of cheese-making dates to 5500 BCE in Kujawy, Poland.
Archaeologists seem to agree that the culture of the early Neolithic is relatively homogeneous, compared both to the late Mesolithic and the later Neolithic. The diffusion across Europe, from the Aegean to Britain, took about 2,500 years (6500 BCE - 4000 BCE). The Baltic region was penetrated a bit later, around 3500 BCE, and there was also a delay in settling the Pannonian plain. In general, colonization shows a "saltatory" pattern, as the Neolithic advanced from one patch of fertile alluvial soil to another, bypassing mountainous areas. Analysis of radiocarbon dates show clearly that Mesolithic and Neolithic populations lived side by side for as much as a millennium in many parts of Europe, especially in the Iberian peninsula and along the Atlantic coast.
With some exceptions, population levels rose rapidly at the beginning of the Neolithic until they reached the carrying capacity. This was followed by a population crash of "enormous magnitude" after 5000 BCE, with levels remaining low during the next 1,500 years. Populations began to rise after 3500 BCE, with further dips and rises occurring between 3000 and 2500 BCE but varying in date between regions. A study of twelve European regions found most experienced boom and bust patterns and suggested an "endogenous, not climatic cause."
Archaeologists agree that the technologies associated with agriculture originated in the Levant/Near East and then spread into Europe. However, debate exists whether this resulted from an active migratory process from the Near East, or merely due to cultural contact and trade between Europeans and Near Easterners. Currently, three models summarize the proposed pattern of spread:
- Replacement model: posits that there was a significant migration of farmers from the Fertile Crescent into Europe. Given their technological advantages, they would have displaced or absorbed the less numerous hunter-gathering populace. Thus, modern Europeans are primarily descended from these Neolithic farmers.
- Cultural diffusion: in contrast, this model supposes that agriculture reached Europe by way of a flow of ideas and trade between the Mesolithic European population and Anatolian farmers. There was no net increase in migration during this process, and therefore, modern Europeans are descended from the "original" Palaeolithic hunter-gatherers.
- Pioneer model: recognises that models 1) and 2) above may represent false dichotomies. This model postulates that there was an initial, small scale migration of farmers from the Near East to certain regions of Europe. They might have enjoyed localized demographic expansions due to social advantages. The subsequent spread of farming technologies throughout the rest of Europe was then carried out by Mesolithic Europeans who acquired new skill through trade and cultural interaction.
Genetic studies have been utilised in the study of pre-historic population movements. On the whole, scientists agree that there is evidence for a migration during the Neolithic. However, they cannot agree on the extent of this movement. The conclusions of studies appear to be 'operator dependent'. That is, results vary depending on what underlying mutation rates are assumed, and conclusions are drawn from how the authors 'envisage' their results fit with known archaeological and historic processes. Consequently, such studies must be interpreted with caution.
Perhaps the first scholar to posit a large-scale Neolithic migration, based on genetic evidence, was Luigi Luca Cavalli-Sforza. By applying principal component analysis to data from "classical genetic markers" (protein polymorphisms from ABO blood groups, HLA loci, immunoglobulins, etc.), Cavalli-Sforza discovered interesting clues about the genetic makeup of Europeans. Although being very genetically homogeneous, several patterns did exist. The most important one was a north-western to south-eastern cline with a Near Eastern focus. Accounting for 28% of the overall genetic diversity in the European samples in his study, he attributed the cline to the spread of agriculture from the Middle East c. 10,000 to 6,000 years ago.
Cavalli-Sforza's explanation of demic diffusions stipulated that the clines were due to the population expansion of neolithic farmers into a scarcely populated, hunter-gathering Europe, with little initial admixture between agriculturalists and foragers. The predicted route for this spread would have been from Anatolia to central Europe via the Balkans. However, given that the time depths of such patterns are not known, "associating them with particular demographic events is usually speculative". Apart from a demic Neolithic migration, the clines may also be compatible with other demographic scenarios (Barbujani and Bartorelle 2001), such as the initial Palaeolithic expansion, the Mesolithic (post-glacial) re-expansions or later (historic) colonizations.
Studies using direct DNA evidence have produced varying results. A notable proponent of Cavalli-Sforza's demic diffusion scenario is Chikhi. In his 1998 study, utilising polymorphic loci from seven hypervariable autosomal DNA loci, an autocorrelation analysis produced a clinal pattern closely matching that in Cavalli-Sforza’s study. He calculated that the separation times were no older than 10,000 years. "The simplest interpretation of these results is that the current nuclear gene pool largely reflects the westward and northward expansion of a Neolithic group".
Although the above studies propounded a 'significant' Neolithic genetic contribution, they did not quantify the exact magnitude of the genetic contribution. Dupanloup performed an admixture analysis based on several autosomal loci, mtDNA and NRY haplogroup frequencies. The study was based on the assumption that Basques were modern representatives of Palaeolithic hunter-gatherers’ gene pool, and Near Eastern peoples were a proxy population for Neolithic farmers. Subsequently, they used admixture analysis to estimate the likely components of the contemporary European gene pool contributed by the two parental populations whose members hybridized at a certain moment in the past. The study suggested that the greatest Near Eastern admixture occurs in the Balkans (~80%) and Southern Italy (~60%), whilst it is least in peoples of the British Isles (estimating only a 20% contribution). The authors concluded that the Neolithic shift to agriculture entailed major population dispersal from the Near East.
Results derived from analysis of the non-recombining portion of the Y- chromosomes (NRY) produced, at least initially, similar gradients to the classic demic diffusion hypothesis. Two significant studies were Semino 2000 and Rosser 2000, which identified haplogroups J2 and E1b1b (formerly E3b) as the putative genetic signatures of migrating Neolithic farmers from Anatolia, and therefore represent the Y-chromosomal components of a Neolithic demic diffusion.
This association was strengthened when King and Underhill (2002) found that there was a significant correlation between the distribution of Hg J2 and Neolithic painted pottery in European and Mediterranean sites. However, studies of the ancient Y-DNA from the earlier Neolithic cave burials of Cardium pottery culture shows they were mainly haplogroup G2a. These 'Neolithic lineages' accounted for 22% of the total European Y chromosome gene pool, and were predominantly found in Mediterranean regions of Europe (Greece, Italy, southeastern Bulgaria, southeastern Iberia).
Y-DNA based studies
Later Y-DNA based studies, exploiting an increased understanding of the phylogenetic relationships, performing micro-regional haplogroup frequency analysis, reveal a more complicated demographic history. The studies suggest that "the large-scale clinal patterns of Hg E and Hg J reflect a mosaic of numerous small-scale, more regional population movements, replacements, and subsequent expansions overlying previous ranges". Rather than a single, large-scale 'wave of advance' from the Near East, the apparent Hg J2 cline is produced by distinct populations movements emanating from different part of the Aegean and Near East, over a period stretching from the Neolithic to the Classical Period. Similarly, haplogroup E1b1b was also thought to have been introduced into the Balkans by Near Eastern agriculturalists.
However, Cruciani et al. (2007) recently discovered that the large majority of haplogroup E1b1b lineages in Europe are represented by the sub-clade E1b1b1a2- V13, which is rare outside Europe. Cruciani, Battaglia and King all predict that V13 expanded from the Balkans. However, there has been no consensus as to exact timing of this expansion (King and Battalia favour a neolithic expansion, possibly coinciding with the adoption of farming by indigenous Balkaners, whilst Cruciani favours a Bronze Age expansion), nor as to where V13 actually arose (but point to somewhere in the southern Balkans or Anatolia)
Overall, Y-chromosome data seems to support the "Pioneer model", whereby heterogeneous groups of Neolithic farmers colonized selected areas of southern Europe via a primarily maritime route. Subsequent expansion of agriculture was facilitated by the adoption of its methods by indigenous Europeans, a process especially prominent in the Balkans.
The data from mtDNA is also interesting. European mtDNA haplogroup frequencies show little, if any, geographic patterning, a result attributed to different molecular properties of mtDNA, as well as different migratory practices between females and males (Semino 2000). The vast majority of mtDNA lineages (60–70%) have been dated to have emerged either in the Mesolithic or Palaeolithic. whereas only 20% of mitochondrial lineages are "Neolithic". However, this conclusion has been questioned. Any undetected heterogeneity in the founder population would result in an overestimation in the age of the current population's molecular age. If this is true, then Europe could have been populated far more recently, e.g. during the Neolithic, by a more diverse founding population.
As Chikhi states: "We argue that many mitochondrial lineages whose origin has been traced back to the Palaeolithic period probably reached Europe at a later time". However, Richards et al. (2000) maintain these findings even when founding population heterogeneity is considered. In one such study, Wolfgang Haak extracted ancient mtDNA from what they present as early European farmers from the Linear Pottery Culture in central Europe. The bodies contained a 25% frequency of mtDNA N1a, a haplogroup which they assumed to be linked to the Neolithic. Today the frequency of this haplogroup is a mere 0.2%. Haak presented this as supportive evidence for a Palaeolithic European ancestry.
Formerly there had been much debate about whether the westerly spread of agriculture from the Near East was driven by farmers actually migrating, or by the transfer of ideas and technologies to indigenous hunter-gatherers.
However, in 2010, researchers have studied the genetic diversity of modern populations to throw light on the processes involved in these ancient events. The new study, funded by the Wellcome Trust, examines the diversity of the Y chromosome. Mark Jobling, who led the research, said: "We focused on the commonest Y-chromosome lineage in Europe, carried by about 110 million men, it follows a gradient from south-east to north-west, reaching almost 100% frequency in Ireland. We looked at how the lineage is distributed, how diverse it is in different parts of Europe, and how old it is." The results suggested that the lineage R1b1b2 (R-M269), like E1b1b or J lineages, spread together with farming from the Near East. Prior archaeological and metrological studies had arrived at similar conclusions in support of the migrationist model.
Patricia Balaresque, first author of the study, added: "In total, this means that more than 80% of European Y chromosomes descend from incoming farmers. In contrast, most maternal genetic lineages seem to descend from hunter-gatherers. To us, this suggests a reproductive advantage for farming males over indigenous hunter-gatherer males during the switch from hunting and gathering, to farming".
However, in 2011, a study  has argued there to be serious flaws in the above proposed model, denouncing the perceived overgeneralization inherit in the studies of Baleresque 2010. Furthermore, Busby et al. 2012 point out "For this haplogroup to be so ubiquitous, the population carrying R1b-S127 would have displaced most of the populations present in western Europe after the Neolithic agricultural transition". Clearly common sense dictates that this did not happen. Also they go on to show that within the European specific R1b-M269 sub-lineage, defined by SNP S127, there exists distinct sub-haplogroups and at this level there exists several "geographically localized pockets, with individual R1b-M269 sub- haplogroups dominating". Their conclusions were that it is likely that R1b-S127 was already present in native European populations and grew into several geographically distinct sub-lineages across Europe before Neolithic expansion occurred.
In 2015, a thorough study by Haak et al.about ancient DNA, concluded, however, that both R1a and R1b very likely spread into Europe from the Pontic-Caspian steppe after 3,000 BCE. As the authors noticed, there was a paucity of haplogroup R1b (or any other variant of R1) in European population samples predating the Bronze Age, with only one of the 70 individuals from Mesolithic and Neolithic Europe belonging to haplogroup R1. Among the analyzed male samples taken from Yamna culture sites, however, all possessed haplogroup R1b. Analysis of modern Europeans' autosomal DNA also gives support to a large population displacement from the steppe into Europe.
There is no direct evidence of the languages spoken in the Neolithic. Some proponents of paleolinguistics attempt to extend the methods of historical linguistics to the Stone Age, but this has little academic support. Criticising scenarios which envision for the Neolithic only a small number of language families spread over huge areas of Europe (as in modern times), Donald Ringe has argued on general principles of language geography (as concerns "tribal", pre-state societies), and the scant remains of (apparently indigenous) non-Indo-European languages attested in ancient inscriptions, that Neolithic Europe must have been a place of great linguistic diversity, with many language families with no recoverable linguistic links to each other, much like western North America prior to European colonisation.
Discussion of hypothetical languages spoken in the European Neolithic is divided into two topics, Indo-European languages and "Pre-Indo-European" languages.
Early Indo-European languages are usually assumed to have reached the Danubian (and maybe Central) Europe in the Chalcolithic or early Bronze Age, e.g. with the Corded Ware or Beaker cultures (see also Kurgan hypothesis for related discussions). The Anatolian hypothesis postulates arrival of Indo-European languages with the early Neolithic. Old European hydronymy is taken by Hans Krahe to be the oldest reflection of the early presence of Indo-European in Europe.
Theories of "Pre-Indo-European" languages in Europe are built on scant evidence. The Basque language is the best candidate for a descendant of such a language, but since Basque is a language isolate, there is no comparative evidence to build upon. Theo Vennemann nevertheless postulates a "Vasconic" family, which he supposes had co-existed with an "Atlantic" or "Semitidic" (i. e., para-Semitic) group. Another candidate is a Tyrrhenian family which would have given rise to Etruscan and Raetic in the Iron Age, and possibly also Aegean languages such as Minoan or Pelasgian in the Bronze Age.
In the north, a similar scenario to Indo-European is thought to have occurred with Uralic languages expanding in from the east. In particular, while the Sami languages of the indigenous Sami people belong in the Uralic family, they show considerable substrate influence, thought to represent one or more extinct original languages. The Sami are estimated to have adopted a Uralic language less than 2,500 years ago. Some traces of indigenous languages of the Baltic area have been suspected in the Finnic languages as well, but these are much more modest. There are early loanwords from unidentified non-IE languages in other Uralic languages of Europe as well.
List of cultures and sites
- Megalithic culture (8th to 2nd millennia)
- Early Neolithic
- Middle Neolithic
- Vinča culture (6th to 3rd millennia)
- Linear Ceramic culture (6th to 5th millennia)
- Cardium Pottery Culture (Mediterranean coast, 7th to 4th millennia)
- Pit–Comb Ware culture, a.k.a. Comb Ceramic culture (Northeast Europe, 6th to 3rd millennia)
- Pre-Cucuteni culture (Moldova, 5th millennium)
- Ertebølle culture (Denmark, 5th to 3rd millennia)
- Cortaillod culture (Switzerland, 4th millennium)
- Hembury culture (Britain, 5th to 4th millennia)
- Windmill Hill culture (Britain, 3rd millennium)
- Pfyn culture (Switzerland, 4th millennium)
- Globular Amphora culture (Central Europe, 4th to 3rd millennia)
- Horgen culture (Switzerland, 4th to 3rd millennia)
- Eneolithic (Chalcolithic)
- Cucuteni-Trypillian culture (Moldova, 5th to 4th millennia)
- Lengyel culture (5th millennium)
- A culture in Central Europe produced monumental arrangements of circular ditches between 4800 BCE and 4600 BCE.
- Varna culture (5th millennium)
- Funnelbeaker culture (4th millennium)
- Baden culture (Central Europe, 4th to 3rd millennia)
- Corded Ware culture, a.k.a. Battle-axe or Single Grave culture (Northern Europe, 3rd millennium)
- Gaudo culture (3rd millennium, early Bronze Age, in Italian)
- Beaker culture (3rd to 2nd millennia, early Bronze Age)
Some Neolithic cultures listed above are known for constructing megaliths. These occur primarily on the Atlantic coast of Europe, but there are also megaliths on western Mediterranean islands.
- c. 5000 BCE: Constructions in Portugal (Évora). Emergence of the Atlantic Neolithic period, the age of agriculture along the fertile shores of Europe.
- c. 4800 BCE: Constructions in Brittany (Barnenez) and Poitou (Bougon).
- c. 4000 BCE: Constructions in Brittany (Carnac), Portugal (Lisbon), Spain (Galicia and Andalusia), France (central and southern), Corsica, England, Wales, Northern Ireland (Banbridge) and elsewhere.
- c. 3700 BCE: Constructions in Ireland (Carrowmore and elsewhere).
- c. 3600 BCE: Constructions in England (Maumbury Rings and Godmanchester), and Malta (Ġgantija and Mnajdra temples).
- c. 3500 BCE: Constructions in Spain (Málaga and Guadiana), Ireland (south-west), France (Arles and the north), north-west and central Italy (Piedmont, Valle d'Aosta, Liguria and Tuscany), insular Italy (Sardinia and Sicily), Malta (and elsewhere in the Mediterranean), Belgium (north-east) and Germany (central and south-west).
- c. 3400 BCE: Constructions in Ireland (Newgrange), Netherlands (north-east), Germany (northern and central) Sweden and Denmark.
- c. 3200 BCE: Constructions in Malta (Ħaġar Qim and Tarxien).
- c. 3000 BCE: Constructions in France (Saumur, Dordogne, Languedoc, Biscay, and the Mediterranean coast), Spain (Los Millares), Belgium (Ardennes), and Orkney, as well as the first henges (circular earthworks) in Britain.
- c. 2800 BCE: Climax of the megalithic Funnel-beaker culture in Denmark, and the construction of the henge at Stonehenge.
- Ammerman & Cavalli-Sforza 1971.
- Renfrew 1987, p. [page needed].
- Bellwood 2004, p. [page needed].
- Anthony 2007, p. [page needed].
- Bellwood 2004, pp. 68–9.
- Bellwood 2004, pp. 74, 118.
- Subbaraman 2012.
- Bellwood 2004, pp. 68–72.
- Shennan & Edinborough 2007.
- Timpson, Adrian; Colledge, Sue (September 2014). "Reconstructing regional population fluctuations in the European Neolithic using radiocarbon dates: a new case-study using an improved method". Journal of Archaeological Science. 52: 549–557. doi:10.1016/j.jas.2014.08.011.<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Richards et al. 1996.
- Cavalli-Sforza 1997.
- Rosser et al. 2000.
- Di Giacomo et al. 2004.
- Chikhi et al. 1998.
- Dupanloup et al. 2004.
- Semino et al. 2000.
- Lacan et al. 2011.
- Semino et al. 2004.
- Battaglia et al. 2008.
- Barbujani, Bertorelle & Chikhi 1998.
- Vandermeer 1975.
- Zvelebil 2009a.
- Zvelebil 2009b.
- Dokládal & Brožek 1961.
- Bar-Yosef 1998.
- Zvelebil 1989.
- Brace et al. 2005.
- Ricaut & Waelkens 2008.
- Balaresque et al. 2010.
- Busby et al. 2011.
- « R1a and R1b are the most common haplogroups in many European populations today, and our results suggest that they spread into Europe from the East after 3,000 BCE. » in Haak; et al. (2015). "Massive migration from the steppe was a source for Indo-European languages in Europe". Nature. 522: 207–211. doi:10.1038/nature14317. Explicit use of et al. in:
- Derenko et al. 2012.
- Ringe 2009.
- Aikio 2004.
- Häkkinen 2012.
- Aikio, Ante (2004). "An essay on substrate studies and the origin of Saami". In Hyvärinen, Irma; Kallio, Petri; Korhonen, Jarmo (eds.). Etymologie, Entlehnungen und Entwicklungen. Mémoires de la Société Néophilologique de Helsinki (in German). 63. Helsinki: Société Néophilologique. pp. 5–34. ISBN 978-951-9040-19-6. Unknown parameter
|trans_title=ignored (help)CS1 maint: ref=harv (link) CS1 maint: unrecognized language (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Ammerman, A. J.; Cavalli-Sforza, L. L. (1971). "Measuring the Rate of Spread of Early Farming in Europe". Man. 6 (4): 674–88. doi:10.2307/2799190. JSTOR 2799190.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Anthony, David W. (2007). The Horse, the Wheel, and Language: How Bronze-Age Riders from the Eurasian Steppes Shaped the Modern World. Princeton University Press. ISBN 0-691-05887-3.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Balaresque, Patricia; Bowden, Georgina R.; Adams, Susan M.; Leung, Ho-Yee; King, Turi E.; Rosser, Zoë H.; Goodwin, Jane; Moisan, Jean-Paul; Richard, Christelle; Millward, Ann; Demaine, Andrew G.; Barbujani, Guido; Previderè, Carlo; Wilson, Ian J.; Tyler-Smith, Chris; Jobling, Mark A. (2010). Penny, David (ed.). "A Predominantly Neolithic Origin for European Paternal Lineages". PLoS Biology. 8 (1): e1000285. doi:10.1371/journal.pbio.1000285. PMC 2799514. PMID 20087410. Unknown parameter
|laysummary=ignored (help)CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Barbujani, Guido; Bertorelle, Giorgio; Chikhi, Lounès (1998). "Evidence for Paleolithic and Neolithic Gene Flow in Europe". The American Journal of Human Genetics. 62 (2): 488–92. doi:10.1086/301719. PMC 1376895. PMID 9463326.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Bar-Yosef, Ofer (1998). "The Natufian culture in the Levant, threshold to the origins of agriculture". Evolutionary Anthropology: Issues, News, and Reviews. 6 (5): 159–77. doi:10.1002/(SICI)1520-6505(1998)6:5<159::AID-EVAN4>3.0.CO;2-7.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Battaglia, Vincenza; Fornarino, Simona; Al-Zahery, Nadia; Olivieri, Anna; Pala, Maria; Myres, Natalie M; King, Roy J; Rootsi, Siiri; Marjanovic, Damir; Primorac, Dragan; Hadziselimovic, Rifat; Vidovic, Stojko; Drobnic, Katia; Durmishi, Naser; Torroni, Antonio; Santachiara-Benerecetti, A Silvana; Underhill, Peter A; Semino, Ornella (2008). "Y-chromosomal evidence of the cultural diffusion of agriculture in southeast Europe". European Journal of Human Genetics. 17 (6): 820–30. doi:10.1038/ejhg.2008.249. PMC 2947100. PMID 19107149.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Bellwood, Peter (2004). First Farmers: The Origins of Agricultural Societies. Blackwell Publishers. ISBN 0-631-20566-7.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Brace, C. Loring; Seguchi, Noriko; Quintyn, Conrad B.; Fox, Sherry C.; Nelson, A. Russell; Manolis, Sotiris K.; Qifeng, Pan (2005). "The questionable contribution of the Neolithic and the Bronze Age to European craniofacial form". Proceedings of the National Academy of Sciences. 103 (1): 242–7. Bibcode:2006PNAS..103..242B. doi:10.1073/pnas.0509801102. JSTOR 30048282. PMC 1325007. PMID 16371462.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Busby, George B. J.; Brisighelli, Francesca; Sánchez-Diz, Paula; Ramos-Luis, Eva; Martinez-Cadenas, Conrado; Thomas, Mark G.; Bradley, Daniel G.; Gusmão, Leonor; Winney, Bruce; Bodmer, Walter; Vennemann, Marielle; Coia, Valentina; Scarnicci, Francesca; Tofanelli, Sergio; Vona, Giuseppe; Ploski, Rafal; Vecchiotti, Carla; Zemunik, Tatijana; Rudan, Igor; Karachanak, Sena; Toncheva, Draga; Anagnostou, Paolo; Ferri, Gianmarco; Rapone, Cesare; Hervig, Tor; Moen, Torolf; Wilson, James F.; Capelli, Cristian (2011). "The peopling of Europe and the cautionary tale of Y chromosome lineage R-M269". Proceedings of the Royal Society B: Biological Sciences. 279 (1730): 884–92. doi:10.1098/rspb.2011.1044. PMC 3259916. PMID 21865258.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Cavalli-Sforza, LL (1997). "Genes, peoples, and languages". Proceedings of the National Academy of Sciences of the United States of America. 94 (15): 7719–24. Bibcode:1997PNAS...94.7719C. doi:10.1073/pnas.94.15.7719. PMC 33682. PMID 9223254.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Chikhi, L.; Destro-Bisol, G.; Bertorelle, G.; Pascali, V.; Barbujani, G. (1998). "Clines of nuclear DNA markers suggest a largely Neolithic ancestry of the European gene pool". Proceedings of the National Academy of Sciences. 95 (15): 9053–8. Bibcode:1998PNAS...95.9053C. doi:10.1073/pnas.95.15.9053. JSTOR 45884. PMC 21201. PMID 9671803.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Cruciani, F.; et al. (2007). "Tracing past human male movements in northern/eastern Africa and western Eurasia: new clues from Y-chromosomal haplogroups E-M78 and J-M12". Molecular Biology and Evolution. 24 (6): 1300–1311. doi:10.1093/molbev/msm049. PMID 17351267. Explicit use of et al. in:
- Derenko, Miroslava; Malyarchuk, Boris; Denisova, Galina; Perkova, Maria; Rogalla, Urszula; Grzybowski, Tomasz; Khusnutdinova, Elza; Dambueva, Irina; Zakharov, Ilia (2012). Kivisild, Toomas (ed.). "Complete Mitochondrial DNA Analysis of Eastern Eurasian Haplogroups Rarely Found in Populations of Northern Asia and Eastern Europe". PLoS ONE. 7 (2): e32179. Bibcode:2012PLoSO...732179D. doi:10.1371/journal.pone.0032179. PMC 3283723. PMID 22363811.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Di Giacomo, F.; Luca, F.; Popa, L. O.; Akar, N.; Anagnou, N.; Banyko, J.; Brdicka, R.; Barbujani, G.; Papola, F.; Ciavarella, G.; Cucci, F.; Di Stasi, L.; Gavrila, L.; Kerimova, M. G.; Kovatchev, D.; Kozlov, A. I.; Loutradis, A.; Mandarino, V.; Mammi', C.; Michalodimitrakis, E. N.; Paoli, G.; Pappa, K. I.; Pedicini, G.; Terrenato, L.; Tofanelli, S.; Malaspina, P.; Novelletto, A. (2004). "Y chromosomal haplogroup J as a signature of the post-neolithic colonization of Europe". Human Genetics. 115 (5): 357–71. doi:10.1007/s00439-004-1168-9. PMID 15322918.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Dokládal, Milan; Brožek, Josef (1961). "Physical Anthropology in Czechoslovakia: Recent Developments". Current Anthropology. 2 (5): 455–77. doi:10.1086/200228. JSTOR 2739787.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Dupanloup, I.; Bertorelle, G; Chikhi, L; Barbujani, G (2004). "Estimating the Impact of Prehistoric Admixture on the Genome of Europeans". Molecular Biology and Evolution. 21 (7): 1361–72. doi:10.1093/molbev/msh135. PMID 15044595.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Häkkinen, Jaakko (2012). "Early contacts between Uralic and Yukaghir" (PDF). Suomalais-Ugrilaisen Seuran Toimituksia − Mémoires de la Société Finno-Ougrienne. Helsinki: Finno-Ugric Society (264): 91–101. Retrieved 13 July 2013.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Lacan, Marie; Keyser, Christine; Ricaut, François-Xavier; Brucato, Nicolas; Tarrus, Josep; Bosch, Angel; Guilaine, Jean; Crubezy, Eric; Ludes, Bertrand (2011). "Ancient DNA suggests the leading role played by men in the Neolithic dissemination". Proceedings of the National Academy of Sciences. 108 (45): 18255–9. Bibcode:2011PNAS..10818255L. doi:10.1073/pnas.1113061108. PMC 3215063. PMID 22042855.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Renfrew, Colin (1987). Archaeology and Language. London: Jonathan Cape. ISBN 0-521-38675-6.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Ricaut, F. X.; Waelkens, M. (2008). "Cranial Discrete Traits in a Byzantine Population and Eastern Mediterranean Population Movements". Human Biology. 80 (5): 535–64. doi:10.3378/1534-6617-80.5.535. PMID 19341322.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Richards, M; Côrte-Real, H; Forster, P; MacAulay, V; Wilkinson-Herbots, H; Demaine, A; Papiha, S; Hedges, R; Bandelt, HJ; Sykes, B (1996). "Paleolithic and neolithic lineages in the European mitochondrial gene pool". American Journal of Human Genetics. 59 (1): 185–203. PMC 1915109. PMID 8659525.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Ringe, Don (January 6, 2009). "The Linguistic Diversity of Aboriginal Europe". Language Log. Mark Liberman. Retrieved 22 September 2011.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Rosser, Zoë H.; Zerjal, Tatiana; Hurles, Matthew E.; Adojaan, Maarja; Alavantic, Dragan; Amorim, António; Amos, William; Armenteros, Manuel; Arroyo, Eduardo; Barbujani, Guido; Beckman, G; Beckman, L; Bertranpetit, J; Bosch, E; Bradley, DG; Brede, G; Cooper, G; Côrte-Real, HB; De Knijff, P; Decorte, R; Dubrova, YE; Evgrafov, O; Gilissen, A; Glisic, S; Gölge, M; Hill, EW; Jeziorowska, A; Kalaydjieva, L; Kayser, M; Kivisild, T (2000). "Y-Chromosomal Diversity in Europe is Clinal and Influenced Primarily by Geography, Rather than by Language". The American Journal of Human Genetics. 67 (6): 1526–43. doi:10.1086/316890. PMC 1287948. PMID 11078479.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Semino, Ornella; Magri, Chiara; Benuzzi, Giorgia; Lin, Alice A.; Al-Zahery, Nadia; Battaglia, Vincenza; MacCioni, Liliana; Triantaphyllidis, Costas; Shen, Peidong; Oefner, Peter J.; Zhivotovsky, Lev A.; King, Roy; Torroni, Antonio; Cavalli-Sforza, L. Luca; Underhill, Peter A.; Santachiara-Benerecetti, A. Silvana (2004). "Origin, Diffusion, and Differentiation of Y-Chromosome Haplogroups E and J: Inferences on the Neolithization of Europe and Later Migratory Events in the Mediterranean Area". The American Journal of Human Genetics. 74 (5): 1023–34. doi:10.1086/386295. PMC 1181965. PMID 15069642.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Semino, O.; Passarino, G; Oefner, PJ; Lin, AA; Arbuzova, S; Beckman, LE; De Benedictis, G; Francalacci, P; Kouvatsi, A; Limborska, S; Marcikiae, M; Mika, A; Mika, B; Primorac, D; Santachiara-Benerecetti, AS; Cavalli-Sforza, LL; Underhill, PA (2000). "The Genetic Legacy of Paleolithic Homo sapiens sapiens in Extant Europeans: A Y Chromosome Perspective". Science. 290 (5494): 1155–9. Bibcode:2000Sci...290.1155S. doi:10.1126/science.290.5494.1155. PMID 11073453.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Shennan, Stephen; Edinborough, Kevan (2007). "Prehistoric population history: From the Late Glacial to the Late Neolithic in Central and Northern Europe". Journal of Archaeological Science. 34 (8): 1339–45. doi:10.1016/j.jas.2006.10.031.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Subbaraman, Nidhi (2012). "Art of cheese-making is 7,500 years old". Nature. doi:10.1038/nature.2012.12020.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Vandermeer, J. (1975). "Interspecific competition: A new approach to the classical theory". Science. 188 (4185): 253–5. doi:10.1126/science.188.4185.253. PMID 1118725. Unknown parameter
|doi_brokendate=ignored (help)CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Zvelebil, Marek (1989). "On the transition to farming in Europe, or what was spreading with the Neolithic: a reply to Ammerman (1989)". Antiquity. 63 (239): 379–83.CS1 maint: ref=harv (link)<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Zvelebil, Marek (2009). "Mesolithic prelude and neolithic revolution". In Zvelebil, Marek (ed.). Hunters in Transition: Mesolithic Societies of Temperate Eurasia and Their Transition to Farming. Cambridge University Press. pp. 5–15. ISBN 978-0-521-10957-4.<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Zvelebil, Marek (2009). "Mesolithic societies and the transition to farming: problems of time, scale and organisation". In Zvelebil, Marek (ed.). Hunters in Transition: Mesolithic Societies of Temperate Eurasia and Their Transition to Farming. Cambridge University Press. pp. 167–88. ISBN 978-0-521-10957-4.<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Bellwood, Peter (2001). "Early Agriculturalist Population Diasporas? Farming, Languages, and Genes". Annual Review of Anthropology. 30: 181–207. doi:10.1146/annurev.anthro.30.1.181. JSTOR 3069214.<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Cavalli-Sforza, Luigi Luca; Menozzi, Paolo; Piazza, Alberto (1994). The History and Geography of Human Genes. Princeton University Press. ISBN 0-691-08750-4.<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Cavalli-Sforza, Luigi Luca (2001). Genes, Peoples, and Languages. Berkeley: University of California Press. ISBN 0-520-22873-1.<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>
- Gimbutas, Marija (1989). The Language of the Goddess. Harper & Row. ISBN 0-06-250356-1.<templatestyles src="Module:Citation/CS1/styles.css"></templatestyles>