Dilong paradoxus

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Dilong
Temporal range: Early Cretaceous, 125–124.6 Ma
250px
Restored skeleton
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: Saurischia
Suborder: Theropoda
Family: <templatestyles src="Noitalic/styles.css"/>Proceratosauridae
Genus: †Dilong
Xu et al., 2004
Species:
† D. paradoxus
Binomial name
Dilong paradoxus
Xu et al., 2004
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Dilong (which means 'emperor dragon') is a genus of small proceratosaurid tyrannosauroid dinosaur.[1] The only species is Dilong paradoxus. It is from the Lower Cretaceous Yixian Formation near Lujiatun, Beipiao, in the western Liaoning province of China. It lived about 125 million years ago.

Discovery

File:Dilong.jpg
Type specimen

It was described by Xu Xing and colleagues in 2004.[2]

The name is derived from the Chinese meaning 'emperor' and 龙 / 龍 lóng meaning 'dragon'. "Di", "emperor", refers to the relationship of this animal to Tyrannosaurus rex, the "king" tyrannosaurid. "Long" is used to name Chinese dinosaurs in much the same way that the Latin -saur(us) is in the West. The specific name, paradoxus, is a Latinisation of the Ancient Greek παράδοξον meaning 'against received wisdom'.

Description

File:Dilong paradoxus size 01.jpg
Size comparison between Dilong and a human.

The type specimen is IVPP 14243 (Institute of Vertebrate Paleontology and Paleoanthropology in Beijing), a nearly complete, semi-articulated, skull and skeleton. Referred material includes IVPP 1242, a nearly complete skull and presacral vertebrae, TNP01109 (Tianjin Museum of Natural History), a partial skull, and IVPP V11579, another skull which may belong to D. paradoxus, or to a related species. The type specimen of Dilong was about 1.6 m (5.2 ft) in length, but it is thought to be a juvenile and may have been over 2 m (6.6 ft) long when fully grown.

Feathers

See also Feathered dinosaurs

Dilong paradoxus had a covering of simple feathers or protofeathers. The feathers were seen in fossilized skin impressions from near the jaw and tail. They are not identical to modern bird feathers, lacking a central shaft and most likely used for warmth, since they could not have enabled flight. Adult tyrannosaurs, found in Alberta and Mongolia have skin impressions which appear to show the pebbly scales typical of other dinosaurs. Xu et al. (2004) speculated that the tyrannosauroids may have had different skin coverings on different parts of their bodies - perhaps mixing scales and feathers. They also speculated that feathers may correlate negatively with body size - that juveniles may have been feathered, then shed the feathers and expressed only scales as the animal became larger and no longer needed insulation to stay warm.

Classification

File:Dilong TJV 50.JPG
Life restoration.

When Dilong was first described, it was considered one of the earliest and most primitive members of Tyrannosauroidea, the group that includes the later, larger tyrannosaurids such as Tyrannosaurus rex. At least one later study, by Turner and colleagues in 2007, reanalyzed the relationships of coelurosaurian dinosaurs, including Dilong, and found that it was not a tyrannosauroid. Rather, they placed Dilong two steps above the tyrannosauroids in their phylogeny; more advanced than Coelurus, but more primitive than the Compsognathidae.[3] However, other studies continued to find Dilong as a tyrannosauroid, and some (such as Carr & Williamson 2010) found Dilong to fall within Tyrannosauroidea, not among the more advanced coelurosaurs.[4]

Below is a cladogram containing most tyrannosauroids by Loewen et al. in 2013.[5]


Tyrannosauroidea
Proceratosauridae


Proceratosaurus bradleyi



Kileskus aristotocus



Guanlong wucaii





Sinotyrannus kazuoensis




Juratyrant langhami



Stokesosaurus clevelandi







Dilong paradoxus




Eotyrannus lengi




Bagaraatan ostromi




Raptorex kriegsteini




Dryptosaurus aquilunguis





Alectrosaurus olseni



Xiongguanlong baimoensis





Appalachiosaurus montgomeriensis





Alioramus altai



Alioramus remotus




Tyrannosauridae











In a 2014 study, Dilong was found to be a proceratosaurid.[1]

See also

References

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  3. Turner, A.H., Pol, D., Clarke, J.A., Erickson, G.M., and Norell, M. (2007). "Supporting online material for: A basal dromaeosaurid and size evolution preceding avian flight". Science, 317: 1378-1381. doi:10.1126/science.1144066 (supplement)
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External links